About the Authors

Stephen B. Hager

stevehager@augustana.edu

Affiliation Department of Biology, Augustana College, Rock Island, Illinois, United States of America

Bradley J. Cosentino

Affiliation Department of Biology, Hobart and William Smith Colleges, Geneva, New York, United States of America

Kelly J. McKay

Affiliation BioEco Research and Monitoring Center, Hampton, Illinois, United States of America

Cathleen Monson

Affiliation Reynolds, Illinois, United States of America

Walt Zuurdeeg

Affiliation Davenport, Iowa, United States of America

Brian Blevins

Affiliation Davenport, Iowa, United States of America

Competing Interests

The authors have the following interests. Mr. McKay is the owner of BioEco Research and Monitoring Center. He conducts consulting work for businesses and agencies requiring ecological assessments of birds. Mr. McKay has never consulted on nor anticipates future business related to the research in this manuscript. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SBH BJC KJM CM WZ BB. Performed the experiments: SBH KJM CM WZ BB. Analyzed the data: SBH BJC. Wrote the paper: SBH BJC.