About the Authors

Qiang Zhang

Affiliation Department of Biochemistry, Microbiology, and Immunology, University of Ottawa, Ottawa, Ontario, Canada

Mir Munir A. Rahim

Affiliation Department of Biochemistry, Microbiology, and Immunology, University of Ottawa, Ottawa, Ontario, Canada

David S. J. Allan

Affiliation Department of Immunology, University of Toronto, Sunnybrook Research Institute, Toronto, Ontario, Canada

Megan M. Tu

Affiliation Department of Biochemistry, Microbiology, and Immunology, University of Ottawa, Ottawa, Ontario, Canada

Simon Belanger

Affiliation Department of Biochemistry, Microbiology, and Immunology, University of Ottawa, Ottawa, Ontario, Canada

Elias Abou-Samra

Affiliation Department of Biochemistry, Microbiology, and Immunology, University of Ottawa, Ottawa, Ontario, Canada

Jaehun Ma

Affiliation Department of Immunology, University of Toronto, Sunnybrook Research Institute, Toronto, Ontario, Canada

Harman S. Sekhon

Affiliation Department of Pathology and Laboratory Medicine, The Ottawa Hospital, Ottawa, Ontario, Canada

Todd Fairhead

Affiliation Kidney Research Centre, Ottawa Hospital Research Institute, Ottawa, Ontario, Canada

Haggag S. Zein

Affiliations Department of Biochemistry, Microbiology, and Immunology, University of Ottawa, Ottawa, Ontario, Canada, Department of Genetics, Cairo University, Giza, Egypt

James R. Carlyle

amakrigi@uottawa.ca (APM); james.carlyle@utoronto.ca (JRC); andersonst@mail.nih.gov (SKA)

Affiliation Department of Immunology, University of Toronto, Sunnybrook Research Institute, Toronto, Ontario, Canada

Stephen K. Anderson

amakrigi@uottawa.ca (APM); james.carlyle@utoronto.ca (JRC); andersonst@mail.nih.gov (SKA)

Affiliation Basic Science Program, SAIC-Frederick Inc., Laboratory of Experimental Immunology, Frederick National Laboratory for Cancer Research, Frederick, Maryland, United States of America

Andrew P. Makrigiannis

amakrigi@uottawa.ca (APM); james.carlyle@utoronto.ca (JRC); andersonst@mail.nih.gov (SKA)

Affiliation Department of Biochemistry, Microbiology, and Immunology, University of Ottawa, Ottawa, Ontario, Canada

Competing Interests

None of the authors have any competing financial interests. Dr. Stephen K. Anderson is employed by SAIC-Frederick Inc. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: APM SKA JRC. Performed the experiments: QZ MMAR MMT SB EAS HSZ JM. Analyzed the data: DA HSS TF SKA JRC APM. Wrote the paper: SKA JRC APM.