About the Authors

Silvia E. Smith

Affiliations School of Medicine, Department of Ophthalmology and Visual Sciences, University of Utah, Salt Lake City, Utah, United States of America, Department of Anthropology, University of Utah, Salt Lake City, Utah, United States of America

Patrice Showers-Corneli

Affiliation Department of Biology, Univerity of Utah, Salt Lake City, Utah, United States of America

Caitlin N. Dardenne

Affiliations School of Medicine, Department of Ophthalmology and Visual Sciences, University of Utah, Salt Lake City, Utah, United States of America, Department of Anthropology, University of Utah, Salt Lake City, Utah, United States of America

Henry H. Harpending

Affiliation Department of Anthropology, University of Utah, Salt Lake City, Utah, United States of America

Darren P. Martin

Affiliation Computational Biology Group, Institute of Infectious Diseases and Molecular Medicine, University of Cape Town, South Africa

Robert G. Beiko

beiko@cs.dal.ca

Affiliation Faculty of Computer Science, Dalhousie University, Halifax, Nova Scotia, Canada

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SES PSC CND DPM RGB. Performed the experiments: SES CND DPM RGB. Wrote the paper: SES CND DPM RGB. Contributed to the theoretical framework and analysis: SES PSC CND HCH DPM RGB.