About the Authors

Shio Kobayashi

Affiliation Asian Natural Environmental Science Center (ANESC), The University of Tokyo, Tokyo, Japan

Daisuke Tsugama

Affiliation Asian Natural Environmental Science Center (ANESC), The University of Tokyo, Tokyo, Japan

Shenkui Liu

Affiliation Alkali Soil Natural Environmental Science Center (ASNESC), Northeast Forestry University, Harbin, People’s Republic of China

Tetsuo Takano

takano@anesc.u-tokyo.ac.jp

Affiliation Asian Natural Environmental Science Center (ANESC), The University of Tokyo, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TT SL SK. Performed the experiments: SK DT. Analyzed the data: SK DT. Wrote the paper: SK.