About the Authors

David S. Gokhin

Affiliation Department of Cell Biology, The Scripps Research Institute, La Jolla, California, United States of America

Roberta B. Nowak

Affiliation Department of Cell Biology, The Scripps Research Institute, La Jolla, California, United States of America

Nancy E. Kim

Affiliation Department of Cell Biology, The Scripps Research Institute, La Jolla, California, United States of America

Ernest E. Arnett

Affiliation Department of Biological Structure, University of Washington, Seattle, Washington, United States of America

Albert C. Chen

Affiliation Department of Bioengineering, University of California San Diego, La Jolla, California, United States of America

Robert L. Sah

Affiliation Department of Bioengineering, University of California San Diego, La Jolla, California, United States of America

John I. Clark

Affiliation Department of Biological Structure, University of Washington, Seattle, Washington, United States of America

Velia M. Fowler

velia@scripps.edu

Affiliation Department of Cell Biology, The Scripps Research Institute, La Jolla, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RLS JIC VMF. Performed the experiments: DSG RBN NEK EEA ACC. Analyzed the data: DSG RBN NEK VMF. Contributed reagents/materials/analysis tools: DSG RBN NEK EEA ACC RLS JIC VMF. Wrote the paper: DSG VMF.