About the Authors

Joanna L. Davies

Contributed equally to this work with: Joanna L. Davies, Jean-Baptiste Cazier

Affiliation Department of Statistics, University of Oxford, Oxford, United Kingdom

Jean-Baptiste Cazier

Contributed equally to this work with: Joanna L. Davies, Jean-Baptiste Cazier

Affiliation Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom

Malcolm G. Dunlop

Affiliation Colon Cancer Genetics Group, Institute of Genetics and Molecular Medicine, University of Edinburgh and Medical Research Council Human Genetics Unit, Western General Hospital, Edinburgh, Scotland, United Kingdom

Richard S. Houlston

Affiliation Section of Cancer Genetics, Institute of Cancer Research, Sutton, Surrey, United Kingdom

Ian P. Tomlinson

These authors also contributed equally to this work.

Affiliations Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom, Oxford Comprehensive Biomedical Research Centre, John Radcliffe Hospital, Headington, Oxford, United Kingdom

Chris C. Holmes

cholmes@stats.well.ox.ac.uk

These authors also contributed equally to this work.

Affiliations Department of Statistics, University of Oxford, Oxford, United Kingdom, Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom, MRC Harwell, Harwell Science and Innovation Campus, Oxfordshire, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JLD JBC CCH. Analyzed the data: JLD JBC. Contributed reagents/materials/analysis tools: JLD JBC RSH MGD IPT CCH. Wrote the paper: JLD JBC IPT CCH.