About the Authors

Lewis L. Chen

lchen2@umc.edu

Affiliation Department of Otolaryngology, Neurobiology and Anatomical Sciences, Ophthalmology, University of Mississippi Medical Center, Jackson, Mississippi, United States of America

Daeyeol Lee

Affiliation Department of Neurobiology, Yale University Medical School, New Haven, Connecticut, United States of America

Kikuro Fukushima

Affiliation Department of Physiology, Hokkaido University School of Medicine, Sapporo, Japan

Junko Fukushima

Affiliation Department of Health Science, Hokkaido University School of Medicine, Sapporo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LC DL KF JF. Performed the experiments: KF JF. Analyzed the data: LC DL. Wrote the paper: LC DL KF JF.