About the Authors

Ben Wielstra, Jan W. Arntzen, Niels Raes
Naturalis Biodiversity Center, Leiden, The Netherlands

Ben Wielstra, Wouter Beukema, Andrew K. Skidmore, Albertus G. Toxopeus
University of Twente, Faculty of Geo-Information Science and Earth Observation – ITC, Enschede, The Netherlands

Corresponding Author


Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BW WB JWA AKS AGT NR. Performed the experiments: BW WB. Analyzed the data: BW WB. Wrote the paper: BW.