About the Authors

Joshua S. Madin
Department of Biological Sciences, Macquarie University, Sydney, Australia

Terry P. Hughes, Sean R. Connolly
ARC Centre of Excellence for Coral Reef Studies, James Cook University, Townsville, Australia

Sean R. Connolly
School of Marine Biology and Tropical Biology, James Cook University, Townsville, Australia

Corresponding Author

joshua.madin@mq.edu.au

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JSM SRC. Performed the experiments: JSM. Analyzed the data: JSM. Contributed reagents/materials/analysis tools: JSM TPH. Wrote the paper: JSM SRC TPH.