About the Authors

Johanna R. Reed

Affiliation Microbiology, Immunology, and Cancer Biology Graduate Program, University of Minnesota, Minneapolis, Minnesota, United States of America

Matthew D. Stone

Affiliation Biochemistry, Molecular Biology, and Biophysics, University of Minnesota, Minneapolis, Minnesota, United States of America

Thomas C. Beadnell

Affiliation Department of Laboratory Medicine and Pathology and Masonic Cancer Center, University of Minnesota, Minneapolis, Minnesota, United States of America

Yungil Ryu

Affiliation Department of Laboratory Medicine and Pathology and Masonic Cancer Center, University of Minnesota, Minneapolis, Minnesota, United States of America

Timothy J. Griffin

Affiliation Biochemistry, Molecular Biology, and Biophysics, University of Minnesota, Minneapolis, Minnesota, United States of America

Kathryn L. Schwertfeger

schwe251@umn.edu

Affiliations Microbiology, Immunology, and Cancer Biology Graduate Program, University of Minnesota, Minneapolis, Minnesota, United States of America, Department of Laboratory Medicine and Pathology and Masonic Cancer Center, University of Minnesota, Minneapolis, Minnesota, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JRR KLS. Performed the experiments: JRR MDS TCB YR. Analyzed the data: JRR MDS. Contributed reagents/materials/analysis tools: TJG. Wrote the paper: JRR MDS TJG KLS.