About the Authors

Koichiro Ichimura

Affiliations Department of Cell Biology, University of Oklahoma Health Science Center, Oklahoma City, Oklahoma, United States of America, Department of Anatomy and Life Structure, Juntendo University School of Medicine, Tokyo, Japan

Ekaterina Bubenshchikova

Affiliation Department of Cell Biology, University of Oklahoma Health Science Center, Oklahoma City, Oklahoma, United States of America

Rebecca Powell

Affiliation Department of Cell Biology, University of Oklahoma Health Science Center, Oklahoma City, Oklahoma, United States of America

Yayoi Fukuyo

Affiliation Department of Cell Biology, University of Oklahoma Health Science Center, Oklahoma City, Oklahoma, United States of America

Tomomi Nakamura

Affiliations Department of Cell Biology, University of Oklahoma Health Science Center, Oklahoma City, Oklahoma, United States of America, Department of Biological Science and Technology, Graduate School of Industrial Science and Technology, Tokyo University of Science, Noda, Chiba, Japan

Uyen Tran

Affiliation Department of Cell Biology, Lerner Research Institute/Cleveland Clinic Foundation, Cleveland, Ohio, United States of America

Shoji Oda

Affiliation Department of Integrated Biosciences, Graduate School of Frontier Sciences, The University of Tokyo, Kashiwa, Chiba, Japan

Minoru Tanaka

Affiliation Laboratory of Molecular Genetics for Reproduction, National Institute for Basic Biology, Aichi, Japan

Oliver Wessely

Affiliation Department of Cell Biology, Lerner Research Institute/Cleveland Clinic Foundation, Cleveland, Ohio, United States of America

Hidetake Kurihara

Affiliation Department of Anatomy and Life Structure, Juntendo University School of Medicine, Tokyo, Japan

Tatsuo Sakai

Affiliation Department of Anatomy and Life Structure, Juntendo University School of Medicine, Tokyo, Japan

Tomoko Obara

tomoko-obara@ouhsc.edu

Affiliation Department of Cell Biology, University of Oklahoma Health Science Center, Oklahoma City, Oklahoma, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KI TO. Performed the experiments: KI EB RP YF TN UT OW TO. Analyzed the data: KI TO OW. Contributed reagents/materials/analysis tools: KI EB SO MT HK TS TO. Wrote the paper: KI OW TO.