About the Authors

Maria V. Tsiper

Affiliation Bindley Bioscience Center at Purdue University Discovery Park, West Lafayette, Indiana, United States of America

Jennifer Sturgis

Affiliation Department of Basic Medical Sciences, Purdue University, West Lafayette, Indiana, United States of America

Larisa V. Avramova

Affiliation Bindley Bioscience Center at Purdue University Discovery Park, West Lafayette, Indiana, United States of America

Shilpa Parakh

Affiliation Bindley Bioscience Center at Purdue University Discovery Park, West Lafayette, Indiana, United States of America

Raymond Fatig

Affiliation Department of Medicinal Chemistry and Molecular Pharmacology, Purdue University, West Lafayette, Indiana, United States of America

Ana Juan-García

Affiliation Department of Basic Medical Sciences, Purdue University, West Lafayette, Indiana, United States of America

Nianyu Li

Affiliation Comparative Biology and Safety Sciences, Amgen Inc, Seattle, Washington, United States of America

Bartek Rajwa

Affiliation Bindley Bioscience Center at Purdue University Discovery Park, West Lafayette, Indiana, United States of America

Padma Narayanan

Affiliation Comparative Biology and Safety Sciences, Amgen Inc, Seattle, Washington, United States of America

C. W. Qualls Jr

Affiliation Comparative Biology and Safety Sciences, Amgen Inc, Seattle, Washington, United States of America

J. Paul Robinson

Affiliations Department of Basic Medical Sciences, Purdue University, West Lafayette, Indiana, United States of America, Weldon School of Biomedical Engineering, Purdue University, West Lafayette, Indiana, United States of America

V. Jo Davisson

davisson@purdue.edu

Affiliation Department of Medicinal Chemistry and Molecular Pharmacology, Purdue University, West Lafayette, Indiana, United States of America

Competing Interests

PN, CWQ and NL are employees of Amgen who provided reagents and a grant for these studies. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: VJD MVT BR. Performed the experiments: ROF LVA SP JS AJG. Analyzed the data: MVT VJD BR. Contributed reagents/materials/analysis tools: JPR CWQ PN NL JS. Wrote the paper: MVT VJD. Revisions: BR PN JPR.