About the Authors

Jonathan J. Sager

Affiliations Department of Neurobiology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America, Department of Neurology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America, Pittsburgh Institute of Neurodegenerative Diseases, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Gonzalo E. Torres

Affiliations Department of Neurobiology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America, Department of Pharmacology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America, Pittsburgh Institute of Neurodegenerative Diseases, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Edward A. Burton

eab25@pitt.edu

Affiliations Department of Neurology, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America, Department of Microbiology and Molecular Genetics, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America, Pittsburgh Institute of Neurodegenerative Diseases, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America, Geriatric Research, Education and Clinical Center, Pittsburgh Veterans Affairs Healthcare System, Pittsburgh, Pennsylvania, United States of America, Department of Neurology, Pittsburgh Veterans Affairs Healthcare System, Pittsburgh, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JJS GET EAB. Performed the experiments: JJS EAB. Analyzed the data: JJS EAB. Contributed reagents/materials/analysis tools: GET EAB. Wrote the paper: JJS GET EAB.