About the Authors

Stephanie C. J. Keating

Affiliation Department of Clinical Studies, Ontario Veterinary College, University of Guelph, Guelph, Ontario, Canada

Aurelie A. Thomas

Affiliation Institute of Neuroscience and Comparative Biology Centre, Newcastle University, Newcastle upon Tyne, United Kingdom

Paul A. Flecknell

Affiliation Institute of Neuroscience and Comparative Biology Centre, Newcastle University, Newcastle upon Tyne, United Kingdom

Matthew C. Leach

Matthew.Leach@Newcastle.ac.uk

Affiliation Institute of Neuroscience and Comparative Biology Centre, Newcastle University, Newcastle upon Tyne, United Kingdom

Competing Interests

The authors have declared that no competing interests exist. The commercial organisation Harlan UK donated the animals used in the study. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MCL AAT PAF. Performed the experiments: MCL AAT MCL. Analyzed the data: MCL SCJK. Wrote, read, edited and approved the final manuscript: SCJK AAT PAF MCL.