About the Authors

Michael J. Twiner

mtwiner@umd.umich.edu

Affiliations Marine Biotoxins Program, National Oceanic and Atmospheric Administration/National Ocean Service, Charleston, South Carolina, United States of America, Department of Natural Sciences, University of Michigan-Dearborn, Dearborn, Michigan, United States of America

Leanne J. Flewelling

Affiliation Florida Fish and Wildlife Conservation Commission, Fish and Wildlife Research Institute, St. Petersburg, Florida, United States of America

Spencer E. Fire

Affiliation Marine Biotoxins Program, National Oceanic and Atmospheric Administration/National Ocean Service, Charleston, South Carolina, United States of America

Sabrina R. Bowen-Stevens

Affiliations National Oceanic and Atmospheric Administration Fisheries, Southeast Fisheries Science Center, Panama City Laboratory, Panama City, Florida, United States of America, National Oceanic and Atmospheric Administration/National Marine Fisheries Service, Southeast Fisheries Science Center, Miami, Florida, United States of America

Joseph K. Gaydos

Affiliation Wildlife Health Center, School of Veterinary Medicine, University of California Davis, Davis, California, United States of America

Christine K. Johnson

Affiliation Wildlife Health Center, School of Veterinary Medicine, University of California Davis, Davis, California, United States of America

Jan H. Landsberg

Affiliation Florida Fish and Wildlife Conservation Commission, Fish and Wildlife Research Institute, St. Petersburg, Florida, United States of America

Tod A. Leighfield

Affiliation Marine Biotoxins Program, National Oceanic and Atmospheric Administration/National Ocean Service, Charleston, South Carolina, United States of America

Blair Mase-Guthrie

Affiliation National Oceanic and Atmospheric Administration/National Marine Fisheries Service, Southeast Fisheries Science Center, Miami, Florida, United States of America

Lori Schwacke

Affiliation National Oceanic and Atmospheric Administration/National Ocean Service, Cooperative Center for Marine Animal Health, Hollings Marine Laboratory, Charleston, South Carolina, United States of America

Frances M. Van Dolah

Affiliation Marine Biotoxins Program, National Oceanic and Atmospheric Administration/National Ocean Service, Charleston, South Carolina, United States of America

Zhihong Wang

Affiliation Marine Biotoxins Program, National Oceanic and Atmospheric Administration/National Ocean Service, Charleston, South Carolina, United States of America

Teresa K. Rowles

Affiliation National Oceanic and Atmospheric Administration Fisheries, Cooperative Center for Marine Animal Health, Silver Spring, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MJT LJF SRB-S JHL TAL BM-G LS FMV TKR. Performed the experiments: MJT LJF SRB-S TAL BM-G ZW. Analyzed the data: MJT LJF SF SRB-S JKG CKJ TAL LS FMV ZW. Wrote the paper: MJT LJF SF.