About the Authors

Hidetaka Matsumoto

Contributed equally to this work with: Hidetaka Matsumoto, Koji Shibasaki

hide-m@showa.gunma-u.ac.jp

Affiliation Department of Ophthalmology, Gunma University Graduate School of Medicine, Maebashi, Gunma, Japan

Koji Shibasaki

Contributed equally to this work with: Hidetaka Matsumoto, Koji Shibasaki

Affiliation Department of Molecular and Cellular Neurobiology, Gunma University Graduate School of Medicine, Maebashi, Gunma, Japan

Motokazu Uchigashima

Affiliation Department of Anatomy, Hokkaido University School of Medicine, Sapporo, Japan

Amane Koizumi

Affiliation Section of Correlative Physiology, National Institute for Physiological Sciences, Okazaki, Japan

Masashi Kurachi

Affiliation Department of Molecular and Cellular Neurobiology, Gunma University Graduate School of Medicine, Maebashi, Gunma, Japan

Yasuhiro Moriwaki

Affiliation Department of Pharmacology, Faculty of Pharmacy, Keio University, Minato-ku, Tokyo, Japan

Hidemi Misawa

Affiliation Department of Pharmacology, Faculty of Pharmacy, Keio University, Minato-ku, Tokyo, Japan

Koichiro Kawashima

Affiliation Department of Molecular Pharmacology, Kitasato University School of Pharmacy, Minato-ku, Tokyo, Japan

Masahiko Watanabe

Affiliation Department of Anatomy, Hokkaido University School of Medicine, Sapporo, Japan

Shoji Kishi

Affiliation Department of Ophthalmology, Gunma University Graduate School of Medicine, Maebashi, Gunma, Japan

Yasuki Ishizaki

Affiliation Department of Molecular and Cellular Neurobiology, Gunma University Graduate School of Medicine, Maebashi, Gunma, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KS SK YI. Performed the experiments: HM KS MU MK. Analyzed the data: HM KS MU AK MW. Contributed reagents/materials/analysis tools: AK YM HM KK MW. Wrote the paper: HM KS.