About the Authors

Emily S. Charlson

Contributed equally to this work with: Emily S. Charlson, Kyle Bittinger

Affiliations Department of Microbiology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America, Pulmonary, Allergy and Critical Care Division, Department of Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Kyle Bittinger

Contributed equally to this work with: Emily S. Charlson, Kyle Bittinger

Affiliation Department of Microbiology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Jun Chen

Affiliation Infectious Disease Division, Department of Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America; Department of Biostatistics and Epidemiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Joshua M. Diamond

Affiliation Pulmonary, Allergy and Critical Care Division, Department of Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Hongzhe Li

Affiliation Infectious Disease Division, Department of Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America; Department of Biostatistics and Epidemiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Ronald G. Collman

collmanr@mail.med.upenn.edu (RGC); bushman@mail.med.upenn.edu (FDB)

Affiliations Department of Microbiology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America, Pulmonary, Allergy and Critical Care Division, Department of Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Frederic D. Bushman

collmanr@mail.med.upenn.edu (RGC); bushman@mail.med.upenn.edu (FDB)

Affiliation Department of Microbiology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ESC KB RGC FDB. Performed the experiments: ESC KB JC. Analyzed the data: ESC KB JC RGC FDB. Contributed reagents/materials/analysis tools: JMD HL. Wrote the paper: ESC KB RGC FDB.