About the Authors

Khai C. Ang

khaichung@gmail.com (KCA); kcheng76@gmail.com (KCC)

Affiliation Department of Experimental Pathology & Jake Gittlen Cancer Research Foundation, Penn State College of Medicine, Hershey, Pennsylvania, United States of America

Mee S. Ngu

Affiliation School of Environmental and Natural Resource Sciences, Faculty of Science and Technology, Universiti Kebangsaan Malaysia, Bangi, Selangor, Malaysia

Katherine P. Reid

Affiliation Department of Experimental Pathology & Jake Gittlen Cancer Research Foundation, Penn State College of Medicine, Hershey, Pennsylvania, United States of America

Mei S. Teh

Affiliation School of Environmental and Natural Resource Sciences, Faculty of Science and Technology, Universiti Kebangsaan Malaysia, Bangi, Selangor, Malaysia

Zamzuraida S. Aida

Affiliation School of Environmental and Natural Resource Sciences, Faculty of Science and Technology, Universiti Kebangsaan Malaysia, Bangi, Selangor, Malaysia

Danny XR. Koh

Affiliation School of Environmental and Natural Resource Sciences, Faculty of Science and Technology, Universiti Kebangsaan Malaysia, Bangi, Selangor, Malaysia

Arthur Berg

Affiliation Department of Biostatistics & Bioinformatics, Penn State College of Medicine, Hershey, Pennsylvania, United States of America

Stephen Oppenheimer

Affiliation Institute of Cognitive and Evolutionary Anthropology, School of Anthropology and Museum Ethnography: University of Oxford, Oxford, United Kingdom

Hood Salleh

Affiliation Academic Heritage Museum, Chancellery Building, Universiti Kebangsaan Malaysia, Bangi, Selangor, Malaysia

Mahani M. Clyde

Affiliation School of Environmental and Natural Resource Sciences, Faculty of Science and Technology, Universiti Kebangsaan Malaysia, Bangi, Selangor, Malaysia

Badrul M. Md-Zain

Affiliation School of Environmental and Natural Resource Sciences, Faculty of Science and Technology, Universiti Kebangsaan Malaysia, Bangi, Selangor, Malaysia

Victor A. Canfield

Affiliation Department of Pharmacology, Penn State College of Medicine, Hershey, Pennsylvania, United States of America

Keith C. Cheng

khaichung@gmail.com (KCA); kcheng76@gmail.com (KCC)

Affiliation Department of Experimental Pathology & Jake Gittlen Cancer Research Foundation, Penn State College of Medicine, Hershey, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KCC KCA VAC. Performed the experiments: KCA MSN MST ZSA DXRK AB. Analyzed the data: KCA KCC VAC AB. Contributed reagents/materials/analysis tools: KCC KCA AB BMMZ MMC HS. Wrote the paper: KCA KCC KPR SO.