About the Authors

Katriina Tiira, Osmo Hakosalo, Lauri Kareinen, Hannes Lohi
Research Programs Unit, Molecular Medicine, Department of Veterinary Biosciences, University of Helsinki, Helsinki, Finland

Katriina Tiira, Osmo Hakosalo, Lauri Kareinen, Hannes Lohi
The Folkhälsan Research Center, Helsinki, Finland

Anne Thomas
Antagene, Animal Genetics Laboratory, Lyon, France

Anna Hielm-Björkman
Department of Equine and Small Animal Medicine, University of Helsinki, Helsinki, Finland

Catherine Escriou
National Veterinary School of Lyon, Lyon, France

Paul Arnold
Program in Genetics and Genomic Biology, Hospital for Sick Children, Toronto, Ontario, Canada

Corresponding Author

hannes.lohi@helsinki.fi

Competing Interests

AT works in a private Antagene company. This does not alter the authors’ adherence to all the PLoS ONE policies on sharing data and materials. The other authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KT HL. Performed the experiments: KT OH LK AT. Analyzed the data: KT OH AH-B. Contributed reagents/materials/analysis tools: HL AT PA CE. Wrote the paper: KT HL.