About the Authors

Matthew Bates

Contributed equally to this work with: Matthew Bates, Justin O’Grady

Affiliations Division of Infection and Immunity, Department of Infection, University College London, London, United Kingdom, University of Zambia and University College London Medical School (UNZA-UCLMS) Research and Training Project, University Teaching Hospital, Lusaka, Zambia

Justin O’Grady

Contributed equally to this work with: Matthew Bates, Justin O’Grady

Affiliations Division of Infection and Immunity, Department of Infection, University College London, London, United Kingdom, University of Zambia and University College London Medical School (UNZA-UCLMS) Research and Training Project, University Teaching Hospital, Lusaka, Zambia

Peter Mwaba

Affiliations University of Zambia and University College London Medical School (UNZA-UCLMS) Research and Training Project, University Teaching Hospital, Lusaka, Zambia, Ministry of Health, Lusaka, Zambia

Lophina Chilukutu

Affiliation University of Zambia and University College London Medical School (UNZA-UCLMS) Research and Training Project, University Teaching Hospital, Lusaka, Zambia

Judith Mzyece

Affiliation University of Zambia and University College London Medical School (UNZA-UCLMS) Research and Training Project, University Teaching Hospital, Lusaka, Zambia

Busiku Cheelo

Affiliation University of Zambia and University College London Medical School (UNZA-UCLMS) Research and Training Project, University Teaching Hospital, Lusaka, Zambia

Moses Chilufya

Affiliation University of Zambia and University College London Medical School (UNZA-UCLMS) Research and Training Project, University Teaching Hospital, Lusaka, Zambia

Lukundo Mukonda

Affiliation University of Zambia and University College London Medical School (UNZA-UCLMS) Research and Training Project, University Teaching Hospital, Lusaka, Zambia

Maxwell Mumba

Affiliation University of Zambia and University College London Medical School (UNZA-UCLMS) Research and Training Project, University Teaching Hospital, Lusaka, Zambia

John Tembo

Affiliation University of Zambia and University College London Medical School (UNZA-UCLMS) Research and Training Project, University Teaching Hospital, Lusaka, Zambia

Mumba Chomba

Affiliation University of Zambia and University College London Medical School (UNZA-UCLMS) Research and Training Project, University Teaching Hospital, Lusaka, Zambia

Nathan Kapata

Affiliations University of Zambia and University College London Medical School (UNZA-UCLMS) Research and Training Project, University Teaching Hospital, Lusaka, Zambia, National Tuberculosis Control Programme, Ministry of Health, Lusaka, Zambia

Andrea Rachow

Affiliation Mbeya Medical Research Programme (MMRP), Mbeya, Tanzania

Petra Clowes

Affiliation Mbeya Medical Research Programme (MMRP), Mbeya, Tanzania

Markus Maeurer

Affiliation Department of Microbiology, Tumour and Cell Biology, Karolinska Institute, Stockholm, Sweden

Michael Hoelscher

Affiliations Mbeya Medical Research Programme (MMRP), Mbeya, Tanzania, Department for Infectious Diseases and Tropical Medicine, Klinikum of the University of Munich, Munich, Germany

Alimuddin Zumla

a.zumla@ucl.ac.uk

Affiliations Division of Infection and Immunity, Department of Infection, University College London, London, United Kingdom, University of Zambia and University College London Medical School (UNZA-UCLMS) Research and Training Project, University Teaching Hospital, Lusaka, Zambia

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AZ MH PM MB JO. Performed the experiments: LC JM BC M. Chilufya LM M. Mumba JT M. Chomba. Analyzed the data: AZ MB JO. Wrote the paper: AZ MB JO PM LC JM BC M. Chilufya LM M. Mumba JT M. Chomba NK AR PC M. Maeurer MH.