About the Authors

Graham C. Smith

graham.smith@fera.gsi.gov.uk

Affiliation The Food and Environment Research Agency, York, North Yorkshire, United Kingdom

Robbie A. McDonald

Affiliation Environment and Sustainability Institute, University of Exeter, Penryn, Cornwall, United Kingdom

David Wilkinson

Affiliation The Food and Environment Research Agency, York, North Yorkshire, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GCS DW. Performed the experiments: DW. Analyzed the data: DW GCS. Wrote the paper: GCS DW RAM. Conceived and designed the experiments: GCS DW. Performed the experiments: DW. Analyzed the data: DW GCS. Wrote the paper: GCS DW RAM.