About the Authors

Stephane Boyer

stephane.boyer@lincoln.ac.nz

Affiliation Department of Ecology, Faculty of Agriculture and Life Sciences, Lincoln University, Lincoln, New Zealand

Samuel D. J. Brown

Affiliation Bio-Protection Research Centre, Lincoln University, Lincoln, New Zealand

Rupert A. Collins

Affiliation Bio-Protection Research Centre, Lincoln University, Lincoln, New Zealand

Robert H. Cruickshank

Affiliation Department of Ecology, Faculty of Agriculture and Life Sciences, Lincoln University, Lincoln, New Zealand

Marie-Caroline Lefort

Affiliation Bio-Protection Research Centre, Lincoln University, Lincoln, New Zealand

Jagoba Malumbres-Olarte

Affiliation Department of Ecology, Faculty of Agriculture and Life Sciences, Lincoln University, Lincoln, New Zealand

Stephen D. Wratten

Affiliation Bio-Protection Research Centre, Lincoln University, Lincoln, New Zealand

Competing Interests

The authors have read the journal's policy and have the following conflicts: This work was partly supported by Solid Energy New Zealand Limited as part of the rehabilitation of the Stockton mine. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SB SDW. Performed the experiments: SB. Analyzed the data: SB. Contributed reagents/materials/analysis tools: SB RHC. Wrote the paper: SB SDJB RAC RHC M-CL JM-O SDW. Wrote the software code for sliding window analysis: SDJB.