About the Authors

Robert W. Tilghman

rt9c@virginia.edu

Affiliation Department of Microbiology, Immunology and Cancer Biology, University of Virginia, Charlottesville, Virginia, United States of America

Edik M. Blais

Affiliation Department of Biomedical Engineering, University of Virginia, Charlottesville, Virginia, United States of America

Catharine R. Cowan

Affiliation Department of Microbiology, Immunology and Cancer Biology, University of Virginia, Charlottesville, Virginia, United States of America

Nicholas E. Sherman

Affiliation Department of Microbiology, Immunology and Cancer Biology, University of Virginia, Charlottesville, Virginia, United States of America

Pablo R. Grigera

Affiliation Department of Microbiology, Immunology and Cancer Biology, University of Virginia, Charlottesville, Virginia, United States of America

Erin D. Jeffery

Affiliation Department of Microbiology, Immunology and Cancer Biology, University of Virginia, Charlottesville, Virginia, United States of America

Jay W. Fox

Affiliation Department of Microbiology, Immunology and Cancer Biology, University of Virginia, Charlottesville, Virginia, United States of America

Brett R. Blackman

Affiliation Department of Biomedical Engineering, University of Virginia, Charlottesville, Virginia, United States of America

Daniel J. Tschumperlin

Affiliation Molecular and Integrative Physiological Sciences, Harvard School of Public Health, Boston, Massachusetts, United States of America

Jason A. Papin

Affiliation Department of Biomedical Engineering, University of Virginia, Charlottesville, Virginia, United States of America

J. Thomas Parsons

Affiliation Department of Microbiology, Immunology and Cancer Biology, University of Virginia, Charlottesville, Virginia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RWT JTP. Performed the experiments: RWT CRC PRG EDJ. Analyzed the data: RWT EMB NES BRB JAP JTP. Contributed reagents/materials/analysis tools: DJT. Wrote the paper: RWT EMB CRC JWF DJT JAP JTP.