About the Authors

Wenheng Zhang

Affiliation Department of Organismic and Evolutionary Biology, Harvard University Herbaria, Cambridge, Massachusetts, United States of America

Elena M. Kramer

Affiliation Department of Organismic and Evolutionary Biology, Harvard University Herbaria, Cambridge, Massachusetts, United States of America

Charles C. Davis

cdavis@oeb.harvard.edu

Affiliation Department of Organismic and Evolutionary Biology, Harvard University Herbaria, Cambridge, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Analyzed the data: WZ EMK CCD. Wrote the paper: WZ EMK CCD. Conceived and designed the research: WZ EMK CCD. Performed the research: WZ.