About the Authors

Sheri L. Bonar

Contributed equally to this work with: Sheri L. Bonar, Susannah D. Brydges, James L. Mueller

Affiliation Division of Bone and Mineral Diseases, Washington University School of Medicine, St. Louis, Missouri, United States of America

Susannah D. Brydges

Contributed equally to this work with: Sheri L. Bonar, Susannah D. Brydges, James L. Mueller

Affiliation Division of Allergy, Immunology, and Rheumatology and Departments of Pediatrics and Medicine, University of California San Diego, La Jolla, California, United States of America

James L. Mueller

Contributed equally to this work with: Sheri L. Bonar, Susannah D. Brydges, James L. Mueller

Affiliation Division of Allergy, Immunology, and Rheumatology and Departments of Pediatrics and Medicine, University of California San Diego, La Jolla, California, United States of America

Matthew D. McGeough

Affiliation Division of Allergy, Immunology, and Rheumatology and Departments of Pediatrics and Medicine, University of California San Diego, La Jolla, California, United States of America

Carla Pena

Affiliation Division of Allergy, Immunology, and Rheumatology and Departments of Pediatrics and Medicine, University of California San Diego, La Jolla, California, United States of America

Debbie Chen

Affiliation Division of Allergy, Immunology, and Rheumatology and Departments of Pediatrics and Medicine, University of California San Diego, La Jolla, California, United States of America

Susan K. Grimston

Affiliation Division of Bone and Mineral Diseases, Washington University School of Medicine, St. Louis, Missouri, United States of America

Cynthia L. Hickman-Brecks

Affiliation Division of Bone and Mineral Diseases, Washington University School of Medicine, St. Louis, Missouri, United States of America

Soumya Ravindran

Affiliation Department of Orthopaedic Surgery, Washington University School of Medicine, St. Louis, Missouri, United States of America

Audrey McAlinden

Affiliation Department of Orthopaedic Surgery, Washington University School of Medicine, St. Louis, Missouri, United States of America

Deborah V. Novack

Affiliation Division of Bone and Mineral Diseases, Washington University School of Medicine, St. Louis, Missouri, United States of America

Daniel L. Kastner

Affiliation Medical Genetics Branch, Department of Health and Human Services/National Institutes of Health/National Health Genome Research Institute, Bethesda, Maryland, United States of America

Roberto Civitelli

Affiliation Division of Bone and Mineral Diseases, Washington University School of Medicine, St. Louis, Missouri, United States of America

Hal M. Hoffman

These authors also contributed equally to this work.

Affiliation Division of Allergy, Immunology, and Rheumatology and Departments of Pediatrics and Medicine, University of California San Diego, La Jolla, California, United States of America

Gabriel Mbalaviele

gmbalavi@DOM.wustl.edu

These authors also contributed equally to this work.

Affiliation Division of Bone and Mineral Diseases, Washington University School of Medicine, St. Louis, Missouri, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GM HMH. Performed the experiments: SLB JLM SDB MDM CP DC SKG CLH SR. Contributed reagents/materials/analysis tools: AM SKG DVN DLK RC. Wrote the paper: GM. Reviewed the manuscript: SDB DVN RC HMH.