About the Authors

Melanie Clapham

melanie.clapham@cumbria.ac.uk

Affiliation Centre for Wildlife Conservation, National School of Forestry, University of Cumbria, Penrith, United Kingdom

Owen T. Nevin

Affiliation Centre for Wildlife Conservation, National School of Forestry, University of Cumbria, Penrith, United Kingdom

Andrew D. Ramsey

Affiliation Centre for Wildlife Conservation, National School of Forestry, University of Cumbria, Penrith, United Kingdom

Frank Rosell

Affiliation Faculty of Art and Sciences, Department of Environmental and Health Studies, Telemark University College, Bø i Telemark, Norway

Competing Interests

The authors have the following competing interest: This work was funded in part by Knight Inlet Lodge, British Columbia, Canada. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MC OTN ADR FR. Performed the experiments: MC OTN. Analyzed the data: MC OTN ADR. Contributed reagents/materials/analysis tools: MC OTN ADR FR. Wrote the paper: MC OTN ADR FR.