About the Authors

William R. Swindell

Affiliation Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

Xianying Xing

Affiliation Department of Dermatology, University of Michigan School of Medicine, Ann Arbor, Michigan, United States of America

Philip E. Stuart

Affiliation Department of Dermatology, University of Michigan School of Medicine, Ann Arbor, Michigan, United States of America

Cynthia S. Chen

Affiliation Department of Dermatology, University of Michigan School of Medicine, Ann Arbor, Michigan, United States of America

Abhishek Aphale

Affiliation Department of Dermatology, University of Michigan School of Medicine, Ann Arbor, Michigan, United States of America

Rajan P. Nair

Affiliation Department of Dermatology, University of Michigan School of Medicine, Ann Arbor, Michigan, United States of America

John J. Voorhees

Affiliation Department of Dermatology, University of Michigan School of Medicine, Ann Arbor, Michigan, United States of America

James T. Elder

Affiliation Department of Dermatology, University of Michigan School of Medicine, Ann Arbor, Michigan, United States of America

Andrew Johnston

Affiliation Department of Dermatology, University of Michigan School of Medicine, Ann Arbor, Michigan, United States of America

Johann E. Gudjonsson

johanng@med.umich.edu

Affiliation Department of Dermatology, University of Michigan School of Medicine, Ann Arbor, Michigan, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WRS PES RPN JJV JTE AJ JEG. Performed the experiments: XX CSC AA AJ JEG. Analyzed the data: WRS PES RPN JTE AJ JEG. Wrote the paper: WRS JTE AJ JEG.