About the Authors

Yufei Wang

Affiliation Mucosal Immunology Unit, King's College London, Guy's Hospital, London, United Kingdom

Trevor Whittall

Affiliation Mucosal Immunology Unit, King's College London, Guy's Hospital, London, United Kingdom

Durdana Rahman

Affiliation Mucosal Immunology Unit, King's College London, Guy's Hospital, London, United Kingdom

Evelien M. Bunnik

Affiliation AMC Medical Research, Amsterdam, The Netherlands

Robert Vaughan

Affiliation Department of Tissue Typing, King's College London, Guy's Hospital, London, United Kingdom

Jørgen Schøller

Affiliation Immudex, Copenhagen, Denmark

Lesley A. Bergmeier

Affiliation Queen Mary, University of London, Barts and The London Schools of Medicine and Dentistry, Centre for Clinical and Diagnostic Oral Sciences, London, United Kingdom

David Montefiori

Affiliation Department of Immunology, Duke University Medical Center, Durham, North Carolina, United States of America

Mahavir Singh

Affiliation Lionex GmbH, Braunschweig, Germany

Hanneke Schuitemaker

Affiliation AMC Medical Research, Amsterdam, The Netherlands

Thomas Lehner

thomas.lehner@kcl.ac.uk

Affiliation Mucosal Immunology Unit, King's College London, Guy's Hospital, London, United Kingdom

Competing Interests

The authors have affiliation with Immunudex (JS) and Lionex Gmbh (MS). This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: TL. Performed the experiments: YW TW DR EMB RV JS LAB DM MS HS. Analyzed the data: TL YW. Wrote the paper: TL YW.