About the Authors

John W. J. Moore

Affiliation Centre for Immunology and Infection, Hull York Medical School and Department of Biology, University of York, Heslington, York, United Kingdom

Lynette Beattie

Affiliation Centre for Immunology and Infection, Hull York Medical School and Department of Biology, University of York, Heslington, York, United Kingdom

Jane E. Dalton

Affiliation Centre for Immunology and Infection, Hull York Medical School and Department of Biology, University of York, Heslington, York, United Kingdom

Benjamin M. J. Owens

Current address: Translational Gastroenterology Unit, Experimental Medicine Division, Nuffield Department of Medicine, University of Oxford, Oxford, United Kingdom

Affiliation Centre for Immunology and Infection, Hull York Medical School and Department of Biology, University of York, Heslington, York, United Kingdom

Asher Maroof

Current address: GlaxoSmithkline Biologicals, Hamilton, Montana, United States of America

Affiliation Centre for Immunology and Infection, Hull York Medical School and Department of Biology, University of York, Heslington, York, United Kingdom

Mark C. Coles

Affiliation Centre for Immunology and Infection, Hull York Medical School and Department of Biology, University of York, Heslington, York, United Kingdom

Paul M. Kaye

paul.kaye@york.ac.uk

Affiliation Centre for Immunology and Infection, Hull York Medical School and Department of Biology, University of York, Heslington, York, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JWJM LB MCC PMK. Performed the experiments: JWJM LB JED BMJO AM. Analyzed the data: JWJM LB JED BMJO AM PMK. Contributed reagents/materials/analysis tools: MCC. Wrote the paper: JWJM PMK. Supervised the study: PMK.