About the Authors

Itsuro Koizumi

itsuro@cris.hokudai.ac.jp

Affiliations Creative Research Institution, Hokkaido University, Sapporo, Hokkaido, Japan, Graduate School of Environmental Earth Science, Hokkaido University, Sapporo, Hokkaido, Japan

Nisikawa Usio

Affiliations Center for Transdisciplinary Research, Niigata University, Niigata, Japan, Center for Toki and Ecological Restoration, Niigata University, Sado, Japan

Tadashi Kawai

Affiliation Fisheries Research Department, Wakkanai Fisheries Research Institute, Wakkanai, Hokkaido, Japan

Noriko Azuma

Affiliation Nodai Bioresource Institute, Tokyo University of Agriculture, Abashiri, Japan

Ryuichi Masuda

Affiliation Department of Natural History Sciences, Faculty of Science, Hokkaido University, Sapporo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: IK NU TK RM. Performed the experiments: IK NU TK NA RM. Analyzed the data: IK. Wrote the paper: IK NU.