About the Authors

Eija H. Seppälä

Affiliations Research Programs Unit, Molecular Medicine, University of Helsinki, Helsinki, Finland, Department of Veterinary Biosciences and Department of Medical Genetics, University of Helsinki, Helsinki, Finland, Folkhälsan Institute of Genetics, Helsinki, Finland

Lotta L. E. Koskinen

Affiliations Research Programs Unit, Molecular Medicine, University of Helsinki, Helsinki, Finland, Department of Veterinary Biosciences and Department of Medical Genetics, University of Helsinki, Helsinki, Finland, Folkhälsan Institute of Genetics, Helsinki, Finland

Christina H. Gulløv

Affiliation Department of Small Animal Clinical Sciences, University of Copenhagen, Copenhagen, Denmark

Päivi Jokinen

Affiliations Research Programs Unit, Molecular Medicine, University of Helsinki, Helsinki, Finland, Department of Veterinary Biosciences and Department of Medical Genetics, University of Helsinki, Helsinki, Finland, Folkhälsan Institute of Genetics, Helsinki, Finland

Peter Karlskov-Mortensen

Affiliation Department of Basic Animal and Veterinary Sciences, University of Copenhagen, Copenhagen, Denmark

Luciana Bergamasco

Affiliation College of Veterinary Medicine, Kansas State University, Manhattan, Kansas, United States of America

Izabella Baranowska Körberg

Affiliation Department of Animal Breeding and Genetics, Swedish University of Agricultural Sciences, Uppsala, Sweden

Sigitas Cizinauskas

Affiliation Referral Animal Neurology Hospital Aisti, Vantaa, Finland

Anita M. Oberbauer

Affiliation Department of Animal Science, University of California Davis, Davis, California, United States of America

Mette Berendt

Affiliation Department of Small Animal Clinical Sciences, University of Copenhagen, Copenhagen, Denmark

Merete Fredholm

Affiliation Department of Basic Animal and Veterinary Sciences, University of Copenhagen, Copenhagen, Denmark

Hannes Lohi

hannes.lohi@helsinki.fi

Affiliations Research Programs Unit, Molecular Medicine, University of Helsinki, Helsinki, Finland, Department of Veterinary Biosciences and Department of Medical Genetics, University of Helsinki, Helsinki, Finland, Folkhälsan Institute of Genetics, Helsinki, Finland

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: EHS LLEK HL SC LB PK-M MF. Performed the experiments: EHS LLEK PK-M LB SC. Analyzed the data: EHS LLEK LB SC PK-M HL. Contributed reagents/materials/analysis tools: HL PJ CHG IBK AMO MB MF. Wrote the paper: EHS LLEK HL. Contributed to the preparation of the paper: EHS LLEK HL CHG PK-M LB IBK AMO MB MF.