About the Authors

Jonghan Kim

Affiliation Department of Genetics and Complex Diseases, Harvard School of Public Health, Boston, Massachusetts, United States of America

Yuan Li

Affiliation Department of Genetics and Complex Diseases, Harvard School of Public Health, Boston, Massachusetts, United States of America

Peter D. Buckett

Affiliation Department of Genetics and Complex Diseases, Harvard School of Public Health, Boston, Massachusetts, United States of America

Mark Böhlke

Affiliation Department of Pharmaceutical Sciences, Massachusetts College of Pharmacy & Health Sciences, Boston, Massachusetts, United States of America

Khristy J. Thompson

Affiliation Department of Genetics and Complex Diseases, Harvard School of Public Health, Boston, Massachusetts, United States of America

Masaya Takahashi

Affiliation Department of Radiology, Beth Israel Deaconess Medical Center, Harvard Institutes of Medicine, Boston, Massachusetts, United States of America

Timothy J. Maher

Affiliation Department of Pharmaceutical Sciences, Massachusetts College of Pharmacy & Health Sciences, Boston, Massachusetts, United States of America

Marianne Wessling-Resnick

wessling@hsph.harvard.edu

Affiliation Department of Genetics and Complex Diseases, Harvard School of Public Health, Boston, Massachusetts, United States of America

Competing Interests

JK, YL and MWR are co-inventors on a provisional patent application being prepared by Harvard University. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JK YL MWR. Performed the experiments: JK YL PDB MB. Analyzed the data: JK YL MWR KJT. Contributed reagents/materials/analysis tools: MT TJM. Wrote the paper: JK MWR.