About the Authors

Richard J. Butler


Affiliations Bayerische Staatssammlung für Paläontologie und Geologie, Munich, Germany, GeoBio-Center, Ludwig-Maximilians-Universität München, Munich, Germany

Laura B. Porro

Affiliation Department of Organismal Biology and Anatomy, University of Chicago, Illinois, United States of America

Peter M. Galton

Affiliations Professor Emeritus, University of Bridgeport, Bridgeport, Connecticut, United States of America, Rio Vista, California, United States of America

Luis M. Chiappe

Affiliation The Dinosaur Institute, Natural History Museum of Los Angeles County, Los Angeles, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RJB LBP PMG LMC. Analyzed the data: RJB LBP PMG. Wrote the paper: RJB LBP PMG LMC. Prepared figures: RJB LBP. Analysed CT data and functional morphology: LBP.