About the Authors

Illiana Meurs

Contributed equally to this work with: Illiana Meurs, Laura Calpe-Berdiel

Affiliations Division of Biopharmaceutics, Leiden/Amsterdam Center for Drug Research, Gorlaeus Laboratories, Leiden University, Leiden, The Netherlands, Current position at Department of Endocrinology and Metabolic Diseases, Leiden University Medical Center, Leiden, The Netherlands

Laura Calpe-Berdiel

Contributed equally to this work with: Illiana Meurs, Laura Calpe-Berdiel

Affiliation Division of Biopharmaceutics, Leiden/Amsterdam Center for Drug Research, Gorlaeus Laboratories, Leiden University, Leiden, The Netherlands

Kim L. L. Habets

Affiliation Division of Biopharmaceutics, Leiden/Amsterdam Center for Drug Research, Gorlaeus Laboratories, Leiden University, Leiden, The Netherlands

Ying Zhao

Affiliation Division of Biopharmaceutics, Leiden/Amsterdam Center for Drug Research, Gorlaeus Laboratories, Leiden University, Leiden, The Netherlands

Suzanne J. A. Korporaal

Affiliation Division of Biopharmaceutics, Leiden/Amsterdam Center for Drug Research, Gorlaeus Laboratories, Leiden University, Leiden, The Netherlands

A. Mieke Mommaas

Affiliation Electron Microscopy Section, Department of Molecular Cell Biology, Leiden University Medical Center, Leiden, The Netherlands

Emmanuelle Josselin

Affiliation Centre d'Immunologie de Marseille Luminy, Institut National de la Santé et de la Recherche Médicale, Université de la Méditerranée, Marseille, France

Reeni B. Hildebrand

Affiliation Division of Biopharmaceutics, Leiden/Amsterdam Center for Drug Research, Gorlaeus Laboratories, Leiden University, Leiden, The Netherlands

Dan Ye

Affiliation Division of Biopharmaceutics, Leiden/Amsterdam Center for Drug Research, Gorlaeus Laboratories, Leiden University, Leiden, The Netherlands

Ruud Out

Affiliation Division of Biopharmaceutics, Leiden/Amsterdam Center for Drug Research, Gorlaeus Laboratories, Leiden University, Leiden, The Netherlands

Johan Kuiper

Affiliation Division of Biopharmaceutics, Leiden/Amsterdam Center for Drug Research, Gorlaeus Laboratories, Leiden University, Leiden, The Netherlands

Theo J. C. Van Berkel

Affiliation Division of Biopharmaceutics, Leiden/Amsterdam Center for Drug Research, Gorlaeus Laboratories, Leiden University, Leiden, The Netherlands

Giovanna Chimini

Affiliation Centre d'Immunologie de Marseille Luminy, Institut National de la Santé et de la Recherche Médicale, Université de la Méditerranée, Marseille, France

Miranda Van Eck

m.eck@lacdr.leidenuniv.nl

Affiliation Division of Biopharmaceutics, Leiden/Amsterdam Center for Drug Research, Gorlaeus Laboratories, Leiden University, Leiden, The Netherlands

Competing Interests

Funding from a commercial source (Top Institute Pharma). This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: IM LC-B GC MVE. Performed the experiments: IM LC-B KLLH YZ DY RO RBH SJAK EJ AMM. Analyzed the data: IM LC-B KLLH AMM. Contributed reagents/materials/analysis tools: IM LC-B KLLH YZ DY RO RBH SJAK EJ AMM. Wrote the paper: IM LC-B MVE.