About the Authors

Seiji Shiota

Affiliations Department of Environmental and Preventive Medicine, Oita University Faculty of Medicine, Hasama-machi, Yufu-City, Oita, Japan, Department of General Medicine, Oita University Faculty of Medicine, Hasama-machi, Yufu-City, Oita, Japan

Masahide Watada

Affiliations Department of Environmental and Preventive Medicine, Oita University Faculty of Medicine, Hasama-machi, Yufu-City, Oita, Japan, Department of General Medicine, Oita University Faculty of Medicine, Hasama-machi, Yufu-City, Oita, Japan

Osamu Matsunari

Affiliations Department of Environmental and Preventive Medicine, Oita University Faculty of Medicine, Hasama-machi, Yufu-City, Oita, Japan, Department of General Medicine, Oita University Faculty of Medicine, Hasama-machi, Yufu-City, Oita, Japan

Shun Iwatani

Affiliation Department of Environmental and Preventive Medicine, Oita University Faculty of Medicine, Hasama-machi, Yufu-City, Oita, Japan

Rumiko Suzuki

Affiliation Department of Environmental and Preventive Medicine, Oita University Faculty of Medicine, Hasama-machi, Yufu-City, Oita, Japan

Yoshio Yamaoka

yyamaoka@oita-u.ac.jp

Affiliations Department of Environmental and Preventive Medicine, Oita University Faculty of Medicine, Hasama-machi, Yufu-City, Oita, Japan, Department of Medicine-Gastroenterology, Baylor College of Medicine and Michael E. DeBakey Veterans Affairs Medical Center, Houston, Texas, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SS YY SI RS. Performed the experiments: SS YY MW OM. Analyzed the data: SS YY. Contributed reagents/materials/analysis tools: SS YY MW OM. Wrote the paper: SS YY.