About the Authors

Caleb Marshall Brown


Affiliation Department of Ecology and Evolutionary Biology, University of Toronto, Toronto, Ontario, Canada

Anthony P. Russell

Affiliation Department of Biological Sciences, University of Calgary, Calgary, Alberta, Canada

Competing Interests

Funding for travel to Mongolia was provided in part by Lubrizol Corp. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: CMB. Performed the experiments: CMB APR. Analyzed the data: CMB APR. Wrote the paper: CMB APR.