About the Authors

Heather M. Fritz

Affiliation Department of Pathology, Microbiology and Immunology, University of California Davis, School of Veterinary Medicine, Davis, California, United States of America

Kerry R. Buchholz

Affiliation Department of Microbiology and Immunology, Stanford University School of Medicine, Stanford, California, United States of America

Xiucui Chen

Affiliation Division of Biostatistics, University of California Davis, School of Medicine, Davis, California, United States of America

Blythe Durbin-Johnson

Affiliation Division of Biostatistics, University of California Davis, School of Medicine, Davis, California, United States of America

David M. Rocke

Affiliation Division of Biostatistics, University of California Davis, School of Medicine, Davis, California, United States of America

Patricia A. Conrad

Contributed equally to this work with: Patricia A. Conrad, John C. Boothroyd

john.boothroyd@stanford.edu (JCB); paconrad@ucdavis.edu (PAC)

Affiliation Department of Pathology, Microbiology and Immunology, University of California Davis, School of Veterinary Medicine, Davis, California, United States of America

John C. Boothroyd

Contributed equally to this work with: Patricia A. Conrad, John C. Boothroyd

john.boothroyd@stanford.edu (JCB); paconrad@ucdavis.edu (PAC)

Affiliation Department of Microbiology and Immunology, Stanford University School of Medicine, Stanford, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HMF KRB PAC JCB. Performed the experiments: HMF KRB. Analyzed the data: HMF XC BDJ DMR. Contributed reagents/materials/analysis tools: PAC JCB. Wrote the paper: HMF PAC JCB.