About the Authors

Yu-Wen Chien

yuwen32@gmail.com

Affiliation Department of Epidemiology, Rollins School of Public Health, Emory University, Atlanta, Georgia, United States of America

Bruce R. Levin

Affiliation Department of Biology, Emory University, Atlanta, Georgia, United States of America

Keith P. Klugman

Affiliation Hubert Department of Global Health, Rollins School of Public Health, Emory University, Atlanta, Georgia, United States of America

Competing Interests

KPK has received research funding unrelated to the current paper from Pfizer Vaccines and has consulted for Pfizer Vaccines, Merck, GSK and Sanofi-Aventis. This does not alter the authors adherence to all the PLoS ONE policies on sharing data and materials. YC and BRL report no conflict of interest.

Author Contributions

Conceived and designed the experiments: YC BRL KPK. Performed the experiments: YC. Analyzed the data: YC BRL KPK. Wrote the paper: YC BRL KPK. Wrote, modified and debugged the Berkeley Madonna program: YC BRL. Provided biological and epidemiological input: YC BRL KPK. Performed the simulations: YC.