About the Authors

Clinton D. Francis

clinton.francis@nescent.org

Current address: National Evolutionary Synthesis Center, Durham, North Carolina, United States of America

Affiliation Ecology and Evolutionary Biology, University of Colorado, Boulder, Colorado, United States of America

Catherine P. Ortega

Affiliation San Juan Institute of Natural and Cultural Resources, Fort Lewis College, Durango, Colorado, United States of America

Alexander Cruz

Affiliation Ecology and Evolutionary Biology, University of Colorado, Boulder, Colorado, United States of America

Competing Interests

The authors have read the journal's policy and have the following conflicts: This study was partially funded by ConocoPhillips and Williams Energy. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: CDF CPO. Performed the experiments: CDF CPO. Analyzed the data: CDF. Wrote the paper: CDF. Acquisition of funding: CDF CPO AC. Revision of manuscript and approval of version: CDF CPO AC.