About the Authors

Weiling Zheng

Affiliation Department of Biological Sciences, National University of Singapore, Singapore, Singapore

Zhengyuan Wang

Affiliation Department of Biological Sciences, National University of Singapore, Singapore, Singapore

John E. Collins

Affiliation Vertebrate Development and Genetics, Wellcome Trust Genome Campus, Wellcome Trust Sanger Institute, Hinxton, Cambridge, United Kingdom

Robert M. Andrews

Affiliation Vertebrate Development and Genetics, Wellcome Trust Genome Campus, Wellcome Trust Sanger Institute, Hinxton, Cambridge, United Kingdom

Derek Stemple

Affiliation Vertebrate Development and Genetics, Wellcome Trust Genome Campus, Wellcome Trust Sanger Institute, Hinxton, Cambridge, United Kingdom

Zhiyuan Gong

dbsgzy@nus.edu.sg

Affiliation Department of Biological Sciences, National University of Singapore, Singapore, Singapore

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZG. Performed the experiments: WZ. Analyzed the data: WZ ZW JEC RMA ZG. Contributed reagents/materials/analysis tools: JEC RMA DS. Wrote the paper: WZ ZG.