About the Authors

Nathan A. Haislip

Affiliation Department of Forestry, Wildlife, and Fisheries, Center for Wildlife Health, University of Tennessee, Knoxville, Tennessee, United States of America

Matthew J. Gray

Affiliation Department of Forestry, Wildlife, and Fisheries, Center for Wildlife Health, University of Tennessee, Knoxville, Tennessee, United States of America

Jason T. Hoverman

jason.hoverman@colorado.edu

Current address: Department of Ecology and Evolutionary Biology, University of Colorado, Boulder, Colorado, United States of America

Affiliation Department of Forestry, Wildlife, and Fisheries, Center for Wildlife Health, University of Tennessee, Knoxville, Tennessee, United States of America

Debra L. Miller

Affiliations Department of Forestry, Wildlife, and Fisheries, Center for Wildlife Health, University of Tennessee, Knoxville, Tennessee, United States of America, Veterinary Diagnostic and Investigational Laboratory, College of Veterinary Medicine, University of Georgia, Tifton, Georgia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JTH MJG. Performed the experiments: JTH NAH. Analyzed the data: JTH NAH MJG. Contributed reagents/materials/analysis tools: DLM. Wrote the paper: JTH NAH MJG DLM.