About the Authors

Jennifer T. Wolstenholme

Affiliation Department of Pharmacology and Toxicology, Virginia Commonwealth University, Richmond, Virginia, United States of America

Jon A. Warner

Affiliation Department of Pharmacology and Toxicology, Virginia Commonwealth University, Richmond, Virginia, United States of America

Maria I. Capparuccini

Affiliation Department of Biostatistics, Virginia Commonwealth University, Richmond, Virginia, United States of America

Kellie J. Archer

Affiliation Department of Biostatistics, Virginia Commonwealth University, Richmond, Virginia, United States of America

Keith L. Shelton

Affiliation Department of Pharmacology and Toxicology, Virginia Commonwealth University, Richmond, Virginia, United States of America

Michael F. Miles

mfmiles@vcu.edu

Affiliations Department of Pharmacology and Toxicology, Virginia Commonwealth University, Richmond, Virginia, United States of America, Department of Neurology, Virginia Commonwealth University, Richmond, Virginia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JTW MFM. Performed the experiments: JTW JAW KLS. Analyzed the data: JTW JAW MIC KJA KLS MFM . Wrote the paper: JTW JAW MFM. Edited the manuscript: JTW JAW MIC KJA KLS MFM.