About the Authors

Tomoki Maekawa

Contributed equally to this work with: Tomoki Maekawa, Naoki Takahashi

Affiliations Center for Transdisciplinary Research, Niigata University, Niigata, Japan, Laboratory of Periodontology and Immunology, Division of Oral Science for Health Promotion, Niigata University Graduate School of Medical and Dental Sciences, Niigata, Japan

Naoki Takahashi

Contributed equally to this work with: Tomoki Maekawa, Naoki Takahashi

Affiliations Center for Transdisciplinary Research, Niigata University, Niigata, Japan, Laboratory of Periodontology and Immunology, Division of Oral Science for Health Promotion, Niigata University Graduate School of Medical and Dental Sciences, Niigata, Japan

Koichi Tabeta

Affiliation Center for Transdisciplinary Research, Niigata University, Niigata, Japan

Yukari Aoki

Affiliations Center for Transdisciplinary Research, Niigata University, Niigata, Japan, Laboratory of Periodontology and Immunology, Division of Oral Science for Health Promotion, Niigata University Graduate School of Medical and Dental Sciences, Niigata, Japan

Hirotaka Miyashita

Affiliations Center for Transdisciplinary Research, Niigata University, Niigata, Japan, Laboratory of Periodontology and Immunology, Division of Oral Science for Health Promotion, Niigata University Graduate School of Medical and Dental Sciences, Niigata, Japan

Sayuri Miyauchi

Affiliations Center for Transdisciplinary Research, Niigata University, Niigata, Japan, Laboratory of Periodontology and Immunology, Division of Oral Science for Health Promotion, Niigata University Graduate School of Medical and Dental Sciences, Niigata, Japan

Haruna Miyazawa

Affiliations Center for Transdisciplinary Research, Niigata University, Niigata, Japan, Laboratory of Periodontology and Immunology, Division of Oral Science for Health Promotion, Niigata University Graduate School of Medical and Dental Sciences, Niigata, Japan

Takako Nakajima

Affiliations Center for Transdisciplinary Research, Niigata University, Niigata, Japan, General Dentistry and Clinical Education Unit, Niigata University Medical and Dental Hospital, Niigata, Japan

Kazuhisa Yamazaki

kaz@dent.niigata-u.ac.jp

Affiliations Center for Transdisciplinary Research, Niigata University, Niigata, Japan, Laboratory of Periodontology and Immunology, Division of Oral Science for Health Promotion, Niigata University Graduate School of Medical and Dental Sciences, Niigata, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KT TN KY. Performed the experiments: TM NT YA H. Miyashita H. Miyazawa SM KT. Analyzed the data: KT TN KY. Contributed reagents/materials/analysis tools: NT YA H. Miyashita SM KT. Wrote the paper: KY.