About the Authors

Maura McGrail

mmcgrail@iastate.edu

Affiliation Department of Genetics, Development and Cell Biology, Iowa State University, Ames, Iowa, United States of America

Julia M. Hatler

Affiliation Department of Genetics, Cell Biology and Development, Masonic Cancer Center, Center for Genome Engineering, University of Minnesota, Minneapolis, Minnesota, United States of America

Xianyan Kuang

Affiliation Department of Genetics, Development and Cell Biology, Iowa State University, Ames, Iowa, United States of America

Hsin-Kai Liao

Affiliation Department of Genetics, Development and Cell Biology, Iowa State University, Ames, Iowa, United States of America

Kishore Nannapaneni

Affiliation Departments of Ophthalmology and Biomedical Engineering, University of Iowa, Iowa City, Iowa, United States of America

Kristin E. Noack Watt

Affiliation Department of Genetics, Development and Cell Biology, Iowa State University, Ames, Iowa, United States of America

Juli D. Uhl

Affiliation Department of Genetics, Development and Cell Biology, Iowa State University, Ames, Iowa, United States of America

David A. Largaespada

Affiliation Department of Genetics, Cell Biology and Development, Masonic Cancer Center, Center for Genome Engineering, University of Minnesota, Minneapolis, Minnesota, United States of America

Erik Vollbrecht

Affiliation Department of Genetics, Development and Cell Biology, Iowa State University, Ames, Iowa, United States of America

Todd E. Scheetz

Affiliation Departments of Ophthalmology and Biomedical Engineering, University of Iowa, Iowa City, Iowa, United States of America

Adam J. Dupuy

Affiliation Department of Anatomy and Cell Biology, University of Iowa, Iowa City, Iowa, United States of America

Jesse M. Hostetter

Affiliation Department of Veterinary Pathology, Iowa State University, Ames, Iowa, United States of America

Jeffrey J. Essner

Affiliation Department of Genetics, Development and Cell Biology, Iowa State University, Ames, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MM JJE. Performed the experiments: MM JJE H-KL KENW JU. Analyzed the data: MM JJE JMH DAL JMH. Contributed reagents/materials/analysis tools: KN TES AJD XK EV. Wrote the paper: MM JJE.