About the Authors

Zi-Qiang Zhou

Contributed equally to this work with: Zi-Qiang Zhou, Chia-Yi Shung

Affiliation Shriners Hospitals for Children Portland, Portland, Oregon, United States of America

Chia-Yi Shung

Contributed equally to this work with: Zi-Qiang Zhou, Chia-Yi Shung

Affiliations Shriners Hospitals for Children Portland, Portland, Oregon, United States of America, Department of Cell and Developmental Biology, Oregon Health and Science University, Portland, Oregon, United States of America

Sara Ota

Affiliation Shriners Hospitals for Children Portland, Portland, Oregon, United States of America

Haruhiko Akiyama

Affiliation Department of Orthopaedics, Kyoto University, Kyoto, Japan

Douglas R. Keene

Affiliation Shriners Hospitals for Children Portland, Portland, Oregon, United States of America

Peter J. Hurlin

pjh@shcc.org

Affiliations Shriners Hospitals for Children Portland, Portland, Oregon, United States of America, Department of Cell and Developmental Biology, Oregon Health and Science University, Portland, Oregon, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PJH Z-QZ C-YS. Performed the experiments: Z-QZ C-YS SO DK. Analyzed the data: PJH Z-QZ C-YS SO. Contributed reagents/materials/analysis tools: HA. Wrote the paper: PJH.