About the Authors

Yukiko Nariai

Affiliation Department of Systems Engineering, Shizuoka University, Hamamatsu, Japan

Saki Hayashi

Affiliations Department of Systems Engineering, Shizuoka University, Hamamatsu, Japan, Faculty of Information Science, Shizuoka University, Hamamatsu, Japan

Satoru Morita

Affiliation Department of Systems Engineering, Shizuoka University, Hamamatsu, Japan

Yoshitaka Umemura

Affiliation Department of Systems Engineering, Shizuoka University, Hamamatsu, Japan

Kei-ichi Tainaka

Affiliation Department of Systems Engineering, Shizuoka University, Hamamatsu, Japan

Teiji Sota

Affiliation Department of Zoology, Kyoto University, Kyoto, Japan

John R. Cooley

Affiliations Department of Systems Engineering, Shizuoka University, Hamamatsu, Japan, Department of Ecology and Evolutionary Biology, University of Connecticut, Storrs, Connecticut, United States of America

Jin Yoshimura

jin@sys.eng.shizuoka.ac.jp

Affiliations Department of Systems Engineering, Shizuoka University, Hamamatsu, Japan, Marine Biosystems Research Center, Chiba University, Kamogawa, Chiba, Japan, Department of Environmental and Forest Biology, State University of New York College of Environmental Science and Forestry, Syracuse, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YN SH SM YU KT JY. Performed the experiments: YN SH SM YU. Analyzed the data: YN SH SM YU TS KT JRC JY. Contributed reagents/materials/analysis tools: KT TS JRC. Wrote the paper: YN JY.