About the Authors

Margaret E. Graham

Affiliation Department of Cellular and Molecular Physiology, The Physiological Laboratory, Institute of Translational Medicine, University of Liverpool, Liverpool, United Kingdom

Gerald R. Prescott

Affiliation Department of Cellular and Molecular Physiology, The Physiological Laboratory, Institute of Translational Medicine, University of Liverpool, Liverpool, United Kingdom

James R. Johnson

Affiliation Department of Cellular and Molecular Physiology, The Physiological Laboratory, Institute of Translational Medicine, University of Liverpool, Liverpool, United Kingdom

Mathew Jones

Affiliation Department of Cellular and Molecular Physiology, The Physiological Laboratory, Institute of Translational Medicine, University of Liverpool, Liverpool, United Kingdom

Alice Walmesley

Affiliation Department of Cellular and Molecular Physiology, The Physiological Laboratory, Institute of Translational Medicine, University of Liverpool, Liverpool, United Kingdom

Lee P. Haynes

Affiliation Department of Cellular and Molecular Physiology, The Physiological Laboratory, Institute of Translational Medicine, University of Liverpool, Liverpool, United Kingdom

Alan Morgan

Affiliation Department of Cellular and Molecular Physiology, The Physiological Laboratory, Institute of Translational Medicine, University of Liverpool, Liverpool, United Kingdom

Robert D. Burgoyne

Affiliation Department of Cellular and Molecular Physiology, The Physiological Laboratory, Institute of Translational Medicine, University of Liverpool, Liverpool, United Kingdom

Jeff W. Barclay

barclayj@liv.ac.uk

Affiliation Department of Cellular and Molecular Physiology, The Physiological Laboratory, Institute of Translational Medicine, University of Liverpool, Liverpool, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MEG GRP JRJ MJ AW AM RDB JWB LPH. Performed the experiments: MEG GRP JRJ MJ AW LPH. Analyzed the data: MEG GRP JRJ MJ AW AM RDB JWB LPH. Wrote the paper: AM RDB JWB.