About the Authors

Koji Tajino

Affiliation Department of Intelligence Science and Technology, Graduate School of Informatics, Kyoto University, Kyoto, Japan

Hiroshi Hosokawa

Affiliation Department of Intelligence Science and Technology, Graduate School of Informatics, Kyoto University, Kyoto, Japan

Shingo Maegawa

Affiliation Department of Intelligence Science and Technology, Graduate School of Informatics, Kyoto University, Kyoto, Japan

Kiyoshi Matsumura

Affiliation Faculty of Information Science and Technology, Osaka Institute of Technology, Hirakata, Osaka, Japan

Ajay Dhaka

Affiliation Department of Biological Structure, University of Washington, Seattle, Washington, United States of America

Shigeo Kobayashi

skoba@i.kyoto-u.ac.jp

Affiliation Department of Intelligence Science and Technology, Graduate School of Informatics, Kyoto University, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KT HH SM KM SK. Performed the experiments: KT HH. Analyzed the data: KT. Contributed reagents/materials/analysis tools: AD. Wrote the paper: KT SK.