About the Authors

Mahito Yamada

Affiliation Department of Neurology, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Masafumi Ihara

ihara@kuhp.kyoto-u.ac.jp

Affiliation Department of Neurology, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Yoko Okamoto

Affiliation Department of Neurology, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Takakuni Maki

Affiliation Department of Neurology, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Kazuo Washida

Affiliation Department of Neurology, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Akihiro Kitamura

Affiliation Department of Neurology, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Yoshiki Hase

Affiliation Department of Neurology, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Hidefumi Ito

Affiliation Department of Neurology, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Keizo Takao

Affiliation Section of Behavior Analysis, Center for Genetic Analysis of Behavior, National Institute for Physiological Sciences, Okazaki, Aichi, Japan

Tsuyoshi Miyakawa

Affiliation Division of Systems Medical Science, Institute for Comprehensive Medical Science, Fujita Health University, Toyoake, Aichi, Japan

Raj N. Kalaria

Affiliation Institute for Ageing and Health, WRC, Campus for Ageing & Vitality, Newcastle University, Newcastle-upon-Tyne, United Kingdom

Hidekazu Tomimoto

Affiliation Department of Neurology, Graduate School of Medicine, Mie University, Tsu, Mie, Japan

Ryosuke Takahashi

Affiliation Department of Neurology, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MY MI KT HT. Performed the experiments: MY YO. Analyzed the data: MY MI YO TM KW AK YH HI KT TM RNK HT RT. Contributed reagents/materials/analysis tools: MY. Wrote the paper: MY MI.