About the Authors

Anne G. Lindgren

Affiliations Department of Molecular Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America, Eli and Edythe Broad Center of Regenerative Medicine and Stem Cell Research, University of California Los Angeles, Los Angeles, California, United States of America

Kyle Natsuhara

Affiliation Department of Molecular Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America

E. Tian

Affiliation Department of Molecular Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America

John J. Vincent

Affiliations Department of Molecular Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America, Molecular Biology Institute, University of California Los Angeles, Los Angeles, California, United States of America, Eli and Edythe Broad Center of Regenerative Medicine and Stem Cell Research, University of California Los Angeles, Los Angeles, California, United States of America

Xinmin Li

Affiliations Jonsson Comprehensive Cancer Center, University of California Los Angeles, Los Angeles, California, United States of America, Department of Pathology and Laboratory Medicine, University of California Los Angeles, Los Angeles, California, United States of America

Jing Jiao

Affiliation Department of Molecular and Medical Pharmacology, University of California Los Angeles, Los Angeles, California, United States of America

Hong Wu

Affiliations Jonsson Comprehensive Cancer Center, University of California Los Angeles, Los Angeles, California, United States of America, Department of Molecular and Medical Pharmacology, University of California Los Angeles, Los Angeles, California, United States of America, Eli and Edythe Broad Center of Regenerative Medicine and Stem Cell Research, University of California Los Angeles, Los Angeles, California, United States of America

Utpal Banerjee

Affiliations Department of Molecular Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America, Molecular Biology Institute, University of California Los Angeles, Los Angeles, California, United States of America, Jonsson Comprehensive Cancer Center, University of California Los Angeles, Los Angeles, California, United States of America, Eli and Edythe Broad Center of Regenerative Medicine and Stem Cell Research, University of California Los Angeles, Los Angeles, California, United States of America

Amander T. Clark

clarka@ucla.edu

Affiliations Department of Molecular Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America, Molecular Biology Institute, University of California Los Angeles, Los Angeles, California, United States of America, Jonsson Comprehensive Cancer Center, University of California Los Angeles, Los Angeles, California, United States of America, Eli and Edythe Broad Center of Regenerative Medicine and Stem Cell Research, University of California Los Angeles, Los Angeles, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ATC. Performed the experiments: AGL KN ET JJV XL. Analyzed the data: AGL XL JJ HW UB ATC. Contributed reagents/materials/analysis tools: HW JJ UB. Wrote the paper: AGL ATC.